His research and publications focus on wetland ecology, plant population ecology, community and
ecosystem ecology, and recently on how wetland vegetations and ecosystems in Yangtze estuary respond to natural and anthropogenic disturbance.
Cao, H. (2021). Tidal marsh dynamics in a changing climate: A study on the mechanisms of pioneer establishment and erosion. PhD Thesis. Utrecht University: Utrecht. 134 pp. https://doi.org/10.33540/922, meer
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